The dead and decaying wood in and from these trees is one of the most wildlife-rich and important habitats in Devon.
Other important microhabitats include small rot holes, loose bark and sap runs.
Arlington Court, Cheryl Duerden
Dead branches, decaying wood, holes, cavities, splits, sap runs and loose bark. Ancient trees with wide and hollow trunks. Features support a wealth of wildlife, particularly moths, beetles, flies, fungi, lichens, nesting birds and roosting bats. A range of flowering plants, trees and shrubs are close by and provide nectar and pollen for the moths, beetles and flies.
Birds: Lesser spotted woodpeckers, pied flycatchers, spotted flycatchers.
Insects: Dartmoor ancient oak wood moths (especially beautiful bark moth and silver-streaked beauty), blue ground beetle, six-spotted longhorn beetle, apple lacebug, veteran tree flies (Madiza britannica and Rhipidia ctenophora).
Lichens and fungi: Veteran tree lichens (fungi to be added).
UK priority habitat: Yes
Statutory irreplaceable habitats: Ancient and veteran trees
Ancient and veteran trees are exceptionally rich in wildlife. Ancient trees are those which have reached a great age for their species. An oak tree might live for around 900 years, while a hawthorn may reach around 150 years. They often have very wide trunks (some large trees can look squat) and many are hollow. Due to their age they support features such as dead and decaying wood, fungi, splits, holes, cavities, sap runs and loose or flaking bark.
Veteran trees share many of the features of ancient trees but can be any age, so all ancient trees are also veteran trees but many veteran trees aren’t yet ancient. Some veteran trees can be relatively young but have developed ancient features due to environmental stress. For example, environmental stress can cause trees to reduce their crown. As a result, large bare branches may extend from a smaller, leafy crown. This resembles the antlers of a stag and so oaks are sometimes called ‘stag’s-head oaks’.
Since all ancient trees can also be defined as veteran trees, on this page the term veteran tree is used for both.
Veteran trees are found across Devon, in woodlands and as individual trees in rural and urban areas. Concentrations can be found in wood pasture and parkland habitats. The Woodland Trust’s ancient tree map shows that the majority are found in the southern half of Devon, particularly in east Devon and south-west of Dartmoor.
Consultation Q: Does the map reflect reality or only trees that have been mapped?
There are famous ancient trees in Devon. A few examples include the King John’s Oak, an 800-year-old oak tree in the medieval deer park at Shute in east Devon and the Meavy (Royal) Oak on Dartmoor, which is thought to be 900 years old.
The features found in veteran trees provide microhabitats for a wealth of wildlife including insects, lichens, fungi, mosses, bats and birds.
A few key features, and the wildlife they support, include:
Holes and crevices: Holes provide shelter and nesting sites for a range of species. Lots of birds use holes for nesting, including woodpeckers, tawny owls, barn owls and nuthatches. Focus Species include lesser spotted woodpeckers, pied flycatchers and spotted flycatchers. Most species of bat (other than horseshoes) will roost in tree holes and tiny crevices, including rare Bechstein’s bats. Bats also use trees for maternity roosts in the summer. Ancient trees with wide trunks, where temperatures remain more stable, provide deeper hibernating sites for bats.
Dead and decaying wood: Dead and decaying wood is one of the most wildlife-rich and important microhabitats in Devon. A group of invertebrates (known as saproxylic invertebrates) depend on dead or decaying wood. Different species need different types of decaying wood found in different parts of the tree.
Larvae of rare beetles (such as the six-spotted longhorn beetle now only found at Watersmeet SSSI) feed on the dry rot typical of the heartwood in old trees. Rare fly larvae prefer to feed on moist, white rot, typical of sap wood and younger veteran trees. A group of rare lichens live on well-lit, dry and exposed deadwood.
Veteran trees in Dartmoor’s woodlands support rare invertebrates. These include the endangered blue ground beetle, which hibernates under bark or in dead wood, and rare moths such as the silver-streaked beauty and beautiful bark moth feed on decaying wood found under bark.
Some of the UK’s largest and most attractive flies are associated with decaying wood. In Devon these include the buff-tailed bumblefly, orange-belted plumehorn and the spectacular orange-sided comb-horn.
Bark: Rare lichens live on well-lit, old bark found on veteran trees, while rare barbastelle bats roost under flaky bark. Bark also provides shelter for invertebrates.
Fungi: Many invertebrates, particularly flies, feed on decomposing fungi found on veteran trees, such as bracket fungi.
Veteran trees are important habitats in orchards. For example, the threatened apple lacebug is only known from old apple trees in Devon and Cornwall.
To add fungi, roots and underground mycorrihizal networks.
To add following consultation to ensure the most up-to-date data is used.
Area and distribution
Ancient and veteran trees are recorded on the Ancient Tree Inventory, which is managed by the Woodland Trust and based on volunteer assessment and recording. It’s not a statutory record and some estimate that only 10% of ancient and veteran trees are included.
Trees recorded in Devon since 2004:
Ancient trees: 131
Veteran trees: 1,168
Notable trees: 777
Designations and protection
Veteran trees are in the wood pasture and parkland SSSIs as contributory features, although none are designated individually. Many will be subject to Tree Preservation Orders but there is no central record. It’s thought that 75% are not subject to formal protection.
Condition
There is evidence that in the last century 50% of veteran trees have been lost in parts of England.
There are no specific records of the overall condition of ancient and veteran trees in Devon.
Between 2010 and 2015, deadwood levels were unfavourable in 80% of native woodland habitat and 73% of non-native woodland habitat. This is the single biggest non-historical reason for woods being in unfavourable condition (National Forest Inventory 2020). To discuss.
Much of the nature value of veteran trees, in particular for fungi and invertebrates, lies in their deadwood (large or small diameter, still on the tree or lying on adjacent ground). Concerns over safety and ‘untidiness’ often lead to deadwood being removed and crowns reduced, usually when this is unnecessary.
Lack of management of veteran trees can also reduce their lifespan.
Veteran trees continue to be felled unnecessarily because they’re believed to be dying when bracket fungi are found growing on them. Across our urban and rural landscapes, brash is burnt in bonfires or chipped rather than being left or piled in a corner where it can rot down naturally.
Veteran trees in woodlands (especially conifer plantations on ancient woodland sites) can be overlooked and overshaded by other trees.
Healthy roots with their associated mycorrhizal fungi are essential for healthy veteran trees. Ploughing and compaction by animals and vehicles can directly damage roots or compact soils, which then harms the roots. Built development and works to install underground services have a similar effect. Retaining or reinstating permeable rooting areas is critical.
Veteran trees are being lost due to diseases such as ash dieback. However, the increase in deadwood as a result diseases can be a strange silver lining. Dutch elm disease has led to the loss of nearly every mature elm in the county, while ash dieback is currently leading to a large increase in decaying wood.
The loss of a veteran tree is a disaster for the species that it hosts if there is no other veteran tree nearby they can disperse to. Many species that rely on veteran trees (especially invertebrates and lichens) are only able to disperse over small distances and now often live in isolated refuges. It’s critical that replacement trees are allowed to grow and mature and that networks of future veteran trees are created in order that species can move as they need.
Ancient trees have a relatively high level of resilience compared to younger trees of the same species. However, there’s a risk that trees growing in the open may be more susceptible to flooding or storm damage, even more so if they’re already compromised. Some trees may not be able to fully recover from frost.
High numbers of deer and grey squirrels can harm veteran trees. Squirrels can damage bark on older trees, which may increase the risk of disease. Deer can reduce new growth on trees and prevent future veteran trees from regenerating.
There is funding available from the Forestry Commission to support the management of veteran trees.
Ancient and veteran trees have huge cultural and historic value. They’re much loved, which provides great opportunities to increase awareness and action for existing trees and ensure that a network of future veteran trees is developed.
Better (healthy and wildlife-rich) and more ancient and veteran trees, with existing veteran trees forming the core of a future veteran tree network that’s growing across Devon.
See Find out more below for detailed guidance and sources of funding and advice.
More detailed actions are set out for the following Focus Species:
Follow the links below to the relevant Focus Species pages as well as to relevant habitat and wider theme pages:
Wildlife will benefit from these actions being implemented across Devon. However actions are prioritised to the following High Opportunity Areas (in bold). See Mapping for the LNRS Viewer and information on how areas have been mapped.
High Opportunity Areas include:
Tree planting and natural regeneration should be avoided in the following High Opportunity Areas:
Advice should be obtained before carrying out tree planting in the following High Opportunity Areas to ensure that it adds to, rather than harms, wildlife:
Important:
Always check that actions will not conflict with statutory requirements (for example, statutory wildlife sites such as SSSIs or scheduled monuments) or harm existing wildlife habitats, the historic environment or landscapes. See the Environmental considerations page for more information.
In line with government policy (Keepers of Time, 2022) all ancient and veteran trees (including their root zones) should be considered as statutory irreplaceable habitats and as High Opportunity Areas in this LNRS, whether or not they’re on the national Ancient Woodland Inventory (used for the LNRS) or the LNRS Viewer.
Case studies and Where to visit will be agreed following discussion through public consultation period and then finalised.
There’s lots of great work going on across Devon for ancient and veteran trees. For example:
The Devon Ancient Tree Forum brings together specialists, enthusiasts and ancient tree owners to share best practice and visit sites across Devon
The Flitton Oak, North Molton (details from Cheryl Duerden), nominated for Tree of the Year in 2023. Halo thinning carried out for overshading beech in order to allow continuity of The Flitton Oak.
National Trust? Veteranisation? Saving Devon’s Treescapes project workshops?
Always follow the Countryside Code and keep to footpaths and sites which are managed for public access. A few examples of places to go are given below, for information on visiting these and other sites see the Explore Devon website.
Ancient and veteran trees can be seen in National Trust parklands at Killerton, Saltram, Arlington and Knightshayes and urban parks such as Devonport in Plymouth and Phear Park in Exmouth.
Add more links and tidy up, and finalise following discussion through public consultation.
The Ancient Tree Forum has lots of useful resources.
The Devon Ancient Tree Forum was established in 2005 and runs trips to sites across the county.
Mapped ancient and veteran trees in Devon are shown on the Woodland Trust’s Ancient Tree Inventory (the inventory as of 2025 is shown on the LNRS Viewer). Please add trees to this map (all additions are visited and verified by a Woodland Trust expert).
The Woodland Trust has lots of information on Ancient trees. They also have a series of guidance notes produced in collaboration with the Ancient Tree Forum. Including:
To discuss with the Woodland Trust:
The Woodland Trust and Ancient Tree Forum have produced guidance on caring for ancient and veteran trees on farms and Ancient and Veteran trees: further guidance on management
There is funding through Government agri-environment schemes for management of veteran trees and wood pasture and parkland.
Veteran Trees – A guide to good management, English Nature
An inventory of Devon’s parklands, wood pasture and veteran tree sites was produced in 2007.
Woodland ecological condition in Britain. 2020. National Forest Inventory. Forestry Commission.
The key Government policy position is set out in Keepers of Time. This states that all ancient woodland and ancient and veteran trees are irreplaceable, regardless of whether they are present on the Ancient Woodland Inventory or Ancient Tree Inventory.
VetCert qualifications for veteran tree management professionals are now available across Europe, in the UK via the Arboricultural Assocation.
Prediction of distribution of veteran trees in the UK (published 2021) – This used modelling and the data in the ATI to learn more about the likely distribution of ancient and veteran trees across the UK and produce a distribution map that highlights ‘hotspots’ likely to support unrecorded ancient trees. Based on this, the most conservative estimate of ancient tree abundance in English wood-pastures was 101,402 trees, representing an increase of more than 2,000 % on the known number of ancient trees in wood pastures. This prediction suggests that most ancient trees in the English landscape are yet to be recorded. Full paper here.
To be discussed during consultation
Nature rich
Long Term: 95% under active management.
Short Term: Maintain the existing resource of known ancient and veteran trees.
More and better connected
Long Term: Expanded as part of 16.5% woodland cover by 2050.
Short term: Part of 3,000 ha’s of new woodland by 2030.
Land managers – especially of wood pasture and parkland sites.
National Trust
Woodland Trust
Devon Ancient Tree Forum
Devon Tree Officers
Protected Landscapes
Arboricultural Association
Natural England (SSSIs, ELM)
